Darwinian Daisyworld model from and Watson & Lovelock (1983), Robertson & Robinson (1998) and Lenton & Lovelock (2001).

Units of time are 40 Myr, giving R&R's span of 10 Gyr.

This model offers many opportunities for modification. Maybe the Black and White populations can mutate into each other? The role of q as a measure of segregation of the B and W populations is interesting, and while this model uses purely stigmergic communication between species, direct interaction would also be possible.

Darwinian Daisyworld model from and Watson & Lovelock (1983), Robertson & Robinson (1998) and Lenton & Lovelock (2001).

Units of time are 40 Myr, giving R&R's span of 10 Gyr.

This model offers many opportunities for modification. Maybe the Black and White populations can mutate into each other? The role of q as a measure of segregation of the B and W populations is interesting, and while this model uses purely stigmergic communication between species, direct interaction would also be possible.

Darwinian Daisyworld model from and Watson & Lovelock (1983), Robertson & Robinson (1998) and Lenton & Lovelock (2001).

Units of time are 40 Myr, giving R&R's span of 10 Gyr.

This model offers many opportunities for modification. Maybe the Black and White populations can mutate into each other? The role of q as a measure of segregation of the B and W populations is interesting, and while this model uses purely stigmergic communication between species, direct interaction would also be possible.

Darwinian Daisyworld model from and Watson & Lovelock (1983), Robertson & Robinson (1998) and Lenton & Lovelock (2001).

Units of time are 40 Myr, giving R&R's span of 10 Gyr.

This model offers many opportunities for modification. Maybe the Black and White populations can mutate into each other? The role of q as a measure of segregation of the B and W populations is interesting, and while this model uses purely stigmergic communication between species, direct interaction would also be possible.

Darwinian Daisyworld model from and Watson & Lovelock (1983), Robertson & Robinson (1998) and Lenton & Lovelock (2001).

Units of time are 40 Myr, giving R&R's span of 10 Gyr.

This model offers many opportunities for modification. Maybe the Black and White populations can mutate into each other? The role of q as a measure of segregation of the B and W populations is interesting, and while this model uses purely stigmergic communication between species, direct interaction would also be possible.

Darwinian Daisyworld model from and Watson & Lovelock (1983), Robertson & Robinson (1998) and Lenton & Lovelock (2001).

Units of time are 40 Myr, giving R&R's span of 10 Gyr.

This model offers many opportunities for modification. Maybe the Black and White populations can mutate into each other? The role of q as a measure of segregation of the B and W populations is interesting, and while this model uses purely stigmergic communication between species, direct interaction would also be possible.

Darwinian Daisyworld model from and Watson & Lovelock (1983), Robertson & Robinson (1998) and Lenton & Lovelock (2001).

Units of time are 40 Myr, giving R&R's span of 10 Gyr.

This model offers many opportunities for modification. Maybe the Black and White populations can mutate into each other? The role of q as a measure of segregation of the B and W populations is interesting, and while this model uses purely stigmergic communication between species, direct interaction would also be possible.

Darwinian Daisyworld model from and Watson & Lovelock (1983), Robertson & Robinson (1998) and Lenton & Lovelock (2001).

Units of time are 40 Myr, giving R&R's span of 10 Gyr.

This model offers many opportunities for modification. Maybe the Black and White populations can mutate into each other? The role of q as a measure of segregation of the B and W populations is interesting, and while this model uses purely stigmergic communication between species, direct interaction would also be possible.

A damped, driven pendulum. The combination of damping and driving provokes a minimal case of chaos. The system is not in itself biological, however its chaotic behaviour arises from competing oscillatory influences - a situation which plays a role in many biological systems.

It is easy to imagine allopatric speciation taking place as populations become confined to physically separate locations (for example Darwin's finches on separate Galapagos islands). It is less easy to imagine how sympatric speciation can occur when the nascent species are physically mixed. Golubitsky and Stewart demonstrate a mathematical model of functional segregation in precisely such completely mixed populations.