Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate   algal  ,   growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp
Simple model to illustrate   algal  ,   growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.

  Biogas, model  as well birefineray option to seperate c02 , chp from bogas model are proposed
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Primary production model with phytoplankton as a state variable, force by light and nutrients. Model expanded to include bivalves.
Primary production model with phytoplankton as a state variable, force by light and nutrients. Model expanded to include bivalves.
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate a simple simulation of the microalgae biomass production, focusing on the dependent variables such as light, nutrients and other factor that is running for a yearly period.  The biomass model uses an example, Phytoplankton growth based on Steele's and Michaelis-Menten equa
Simple model to illustrate a simple simulation of the microalgae biomass production, focusing on the dependent variables such as light, nutrients and other factor that is running for a yearly period.

The biomass model uses an example, Phytoplankton growth based on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Once this is understood, it looks upon the viability of biogas production from the microalgae biomass.


Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


 Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I

Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.

Modèle simple pour illustrer la croissance des huîtres sur la base de la production primaire de phytoplancton comme une variable d'état, forcé par la lumière et les éléments nutritifs, en cours d'exécution pour une période annuelle.

La croissance du phytoplancton sur la base de Steele et équations de Michaelis-Menten), où:

Production primaire = (([Pmax] * [I] / [Iopt] * exp (1 - [I] / [Iopt]) * [S]) / ([K] + [S]))

Pmax: production maximale (d-1)
I: L'énergie lumineuse en profondeur de l'intérêt (Ue m-2 s-1)
Iopt: L'énergie lumineuse à laquelle se produit Pmax (Ue m-2 s-1)
S: concentration des éléments nutritifs (N umol L-1)
KS: Demi constants de saturation en nutriments (N umol L-1).

D'autres développements:
- Les éléments nutritifs comme variable d'état dans le cycle de détritus de phytoplancton et d'huîtres de la biomasse.
- Lumière limitée par la concentration de phytoplancton.
- Effet de la température sur le phytoplancton et la croissance des huîtres.

Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate a simple simulation of the microalgae biomass production, focusing on the dependent variables such as light, nutrients and other factor that is running for a yearly period.  The biomass model uses an example, Phytoplankton growth based on Steele's and Michaelis-Menten equa
Simple model to illustrate a simple simulation of the microalgae biomass production, focusing on the dependent variables such as light, nutrients and other factor that is running for a yearly period.

The biomass model uses an example, Phytoplankton growth based on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Once this is understood, it looks upon the viability of biogas production from the microalgae biomass.


Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate   algal  ,   growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp
Simple model to illustrate   algal  ,   growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.

  Biogas, model  as well birefineray option to seperate c02 , chp from bogas model are proposed
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.