Ecology | Insight Maker

Dynamic simulation modelers are particularly interested in understanding and being able to distinguish between the behavior of stocks and flows that result from internal interactions and those that result from external forces acting on a system. For some time modelers have been particularly interested in internal interactions that result in stable oscillations in the absence of any external forces acting on a system. The model in this last scenario was independently developed by Alfred Lotka (1924) and Vito Volterra (1926). Lotka was interested in understanding internal dynamics that might explain oscillations in moth and butterfly populations and the parasitoids that attack them. Volterra was interested in explaining an increase in coastal populations of predatory fish and a decrease in their prey that was observed during World War I when human fishing pressures on the predator species declined. Both discovered that a relatively simple model is capable of producing the cyclical behaviors they observed. Since that time, several researchers have been able to reproduce the modeling dynamics in simple experimental systems consisting of only predators and prey. It is now generally recognized that the model world that Lotka and Volterra produced is too simple to explain the complexity of most and predator-prey dynamics in nature. And yet, the model significantly advanced our understanding of the critical role of feedback in predator-prey interactions and in feeding relationships that result in community dynamics.The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.

2. The food supply of the predator population depends entirely on the size of the prey population.

3. The rate of change of population is proportional to its size.

4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.

5. Predators have limitless appetite.

As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey

When multiplied out, the prey equation becomes

dx/dt = αx - βxy

The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt = -

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.

Clone of Predator-Prey Model ("Lotka'Volterra")

Sean R Westley

Clone of Bio103 Predator-Prey Model ("Lotka'Volterra")

Tags: Education, Chaos, Ecology, Biology, Population

Thanks to Insight Author: John Petersen

Edits by Andy Long

Everything that follows the dashes was created by John Petersen (or at least came from his Insight model). I just wanted to make a few comments.

We are looking at Hare and Lynx, of course. Clone this insight, and change the names.

Then read the text below, to get acquainted with one of the most important and well-known examples of a simple system of differential equations in all of mathematics.

http://www.nku.edu/~longa/classes/mat375/mathematica/Lotka-Volterra.nb

------------------------------------------------------------

Dynamic simulation modelers are particularly interested in understanding and being able to distinguish between the behavior of stocks and flows that result from internal interactions and those that result from external forces acting on a system.

For some time modelers have been particularly interested in internal interactions that result in stable oscillations in the absence of any external forces acting on a system.

The model in this last scenario was independently developed by Alfred Lotka (1924) and Vito Volterra (1926). Lotka was interested in understanding internal dynamics that might explain oscillations in moth and butterfly populations and the parasitoids that attack them. Volterra was interested in explaining an increase in coastal populations of predatory fish and a decrease in their prey that was observed during World War I when human fishing pressures on the predator species declined.

Both discovered that a relatively simple model is capable of producing the cyclical behaviors they observed.

Since that time, several researchers have been able to reproduce the modeling dynamics in simple experimental systems consisting of only predators and prey. It is now generally recognized that the model world that Lotka and Volterra produced is too simple to explain the complexity of most predator-prey dynamics in nature. And yet, the model significantly advanced our understanding of the critical role of feedback in predator-prey interactions and in feeding relationships that result in community dynamics.

The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.

2. The food supply of the predator population depends entirely on the size of the prey population.

3. The rate of change of population is proportional to its size.

4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.

5. Predators have limitless appetite.

As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey

When multiplied out, the prey equation becomes

dx/dt = αx - βxy

The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt = -

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.

Clone of MAT375 Clone of Bio103 Predator-Prey Model ("Lotka'Volterra")

Jacob Adkins

This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

Experiment with adjusting the initial number of moose and wolves on the island.

Clone of Isle Royale: Predator Prey Interactions

Valerya

Overview

A model which simulates the competition between logging versus adventure tourism (mountain bike ridding) in Derby Tasmania. Simulation borrowed from the Easter Island simulation.

How the model works.

Trees grow, we cut them down because of demand for Timber amd sell the logs.

With mountain bkie visits. This depends on past experience and recommendations. Past experience and recommendations depends on Scenery number of trees compared to visitor and Adventure number of trees and users. Park capacity limits the number of users.

Interesting insights

It seems that high logging does not deter mountain biking. By reducing park capacity, visitor experience and numbers are improved. A major problem is that any success with the mountain bike park leads to an explosion in visitor numbers. Also a high price of timber is needed to balance popularity of the park. It seems also that only a narrow corridor is needed for mountain biking

Clone of Simulation of Derby Mountain biking versus logging

Silong Lou

This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

We incorporate logistic growth into the moose dynamics, and we replace the death flow of the moose with a kill rate modeled from the kill rate data found on the Isle Royale website.

I start with these parameters:
Wolf Death Rate = 0.15
Wolf Birth Rate = 0.0187963
Moose Birth Rate = 0.4
Carrying Capacity = 2000
Initial Moose: 563
Initial Wolves: 20

I used RK-4 with step-size 0.1, from 1959 for 60 years.

The moose birth flow is logistic, MBR*M*(1-M/K)
Moose death flow is Kill Rate (in Moose/Year)
Wolf birth flow is WBR*Kill Rate (in Wolves/Year)
Wolf death flow is WDR*W

Clone of Final Midterm Student version of A More Realistic Model of Isle Royale: Predator Prey Interactions

Matthew Gall

Physical meaning of the equations

The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.

2. The food supply of the predator population depends entirely on the size of the prey population.

3. The rate of change of population is proportional to its size.

4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.

5. Predators have limitless appetite.

As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey

When multiplied out, the prey equation becomes

dx/dt = αx - βxy

The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt = -

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.

Clone of Prey&Predator

Chiara Birk

This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

Experiment with adjusting the initial number of moose and wolves on the island.

Clone of Clone of Isle Royale: Predator Prey Interactions

Valerya

This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

We incorporate logistic growth into the moose dynamics, and we replace the death flow of the moose with a kill rate modeled from the kill rate data found on the Isle Royale website.

I start with these parameters:
Wolf Death Rate = 0.15
Wolf Birth Rate = 0.0187963
Moose Birth Rate = 0.4
Carrying Capacity = 2000
Initial Moose: 563
Initial Wolves: 20

I used RK-4 with step-size 0.1, from 1959 for 60 years.

The moose birth flow is logistic, MBR*M*(1-M/K)
Moose death flow is Kill Rate (in Moose/Year)
Wolf birth flow is WBR*Kill Rate (in Wolves/Year)
Wolf death flow is WDR*W

Clone of Final Midterm Student version of A More Realistic Model of Isle Royale: Predator Prey Interactions

Austin Campbell

This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

We incorporate logistic growth into the moose dynamics, and we replace the death flow of the moose with a kill rate modeled from the kill rate data found on the Isle Royale website.

I start with these parameters:
Wolf Death Rate = 0.15
Wolf Birth Rate = 0.0187963
Moose Birth Rate = 0.4
Carrying Capacity = 2000
Initial Moose: 563
Initial Wolves: 20

I used RK-4 with step-size 0.1, from 1959 for 60 years.

The moose birth flow is logistic, MBR*M*(1-M/K)
Moose death flow is Kill Rate (in Moose/Year)
Wolf birth flow is WBR*Kill Rate (in Wolves/Year)
Wolf death flow is WDR*W

Clone of Jacob Englert MAT 375 Midterm: Model of Isle Royale: Predator Prey Interactions

Urasa Kongdechakul

This simulation shows how plant, deer and wolf populations impact each other in a deciduous forest ecosystem.

Clone of Plant, Deer and Wolf Population Dynamics G-IV Intro

Caroline van Eldik

A basic simulation of the spread of Joro Spiders along the US East Coast.

Joro Spider Expansion North USA

Salma Sokkary

Implications of spraying pesticides to control insects. This is an early version of this model.

Systems Thinking Webinars

Clone of Insect Pest Control

Bechara Assouad

Physical meaning of the equations

The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.

2. The food supply of the predator population depends entirely on the size of the prey population.

3. The rate of change of population is proportional to its size.

4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.

5. Predators have limitless appetite.

As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey

When multiplied out, the prey equation becomes

dx/dt = αx - βxy

The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt = -

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.

Clone of Prey&Predator

Helge Klapper

This is an implementation of the 'Very Simple Ecosystem Model' (VSEM) from R package BayesianTools (Hartig et al. 2019). It consists of three stocks: aboveground carbon in plant biomass, belowground carbon in plant biomass and carbon in soil organic matter.

Reference:

Florian Hartig, Francesco Minunno and Stefan Paul (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools

Clone of Very Simple Ecosystem Model

Keenal Shah

Clone of Bio103 Predator-Prey Model ("Lotka'Volterra")

Tags: Education, Chaos, Ecology, Biology, Population

Thanks to Insight Author: John Petersen

Edits by Andy Long

Everything that follows the dashes was created by John Petersen (or at least came from his Insight model). I just wanted to make a few comments.

We are looking at Hare and Lynx, of course. Clone this insight, and change the names.

Then read the text below, to get acquainted with one of the most important and well-known examples of a simple system of differential equations in all of mathematics.

http://www.nku.edu/~longa/classes/mat375/mathematica/Lotka-Volterra.nb

------------------------------------------------------------

Dynamic simulation modelers are particularly interested in understanding and being able to distinguish between the behavior of stocks and flows that result from internal interactions and those that result from external forces acting on a system.

For some time modelers have been particularly interested in internal interactions that result in stable oscillations in the absence of any external forces acting on a system.

The model in this last scenario was independently developed by Alfred Lotka (1924) and Vito Volterra (1926). Lotka was interested in understanding internal dynamics that might explain oscillations in moth and butterfly populations and the parasitoids that attack them. Volterra was interested in explaining an increase in coastal populations of predatory fish and a decrease in their prey that was observed during World War I when human fishing pressures on the predator species declined.

Both discovered that a relatively simple model is capable of producing the cyclical behaviors they observed.

Since that time, several researchers have been able to reproduce the modeling dynamics in simple experimental systems consisting of only predators and prey. It is now generally recognized that the model world that Lotka and Volterra produced is too simple to explain the complexity of most predator-prey dynamics in nature. And yet, the model significantly advanced our understanding of the critical role of feedback in predator-prey interactions and in feeding relationships that result in community dynamics.

The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.

2. The food supply of the predator population depends entirely on the size of the prey population.

3. The rate of change of population is proportional to its size.

4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.

5. Predators have limitless appetite.

As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey

When multiplied out, the prey equation becomes

dx/dt = αx - βxy

The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt = -

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.

Clone of MAT375 Clone of Bio103 Predator-Prey Model ("Lotka'Volterra")

Jordan

Clone of Bio103 Predator-Prey Model ("Lotka'Volterra")

Tags: Education, Chaos, Ecology, Biology, Population

Thanks to Insight Author: John Petersen

Edits by Andy Long

Everything that follows the dashes was created by John Petersen (or at least came from his Insight model). I just wanted to make a few comments.

We are looking at Hare and Lynx, of course. Clone this insight, and change the names.

Then read the text below, to get acquainted with one of the most important and well-known examples of a simple system of differential equations in all of mathematics.

http://www.nku.edu/~longa/classes/mat375/mathematica/Lotka-Volterra.nb

------------------------------------------------------------

Dynamic simulation modelers are particularly interested in understanding and being able to distinguish between the behavior of stocks and flows that result from internal interactions and those that result from external forces acting on a system.

For some time modelers have been particularly interested in internal interactions that result in stable oscillations in the absence of any external forces acting on a system.

The model in this last scenario was independently developed by Alfred Lotka (1924) and Vito Volterra (1926). Lotka was interested in understanding internal dynamics that might explain oscillations in moth and butterfly populations and the parasitoids that attack them. Volterra was interested in explaining an increase in coastal populations of predatory fish and a decrease in their prey that was observed during World War I when human fishing pressures on the predator species declined.

Both discovered that a relatively simple model is capable of producing the cyclical behaviors they observed.

Since that time, several researchers have been able to reproduce the modeling dynamics in simple experimental systems consisting of only predators and prey. It is now generally recognized that the model world that Lotka and Volterra produced is too simple to explain the complexity of most predator-prey dynamics in nature. And yet, the model significantly advanced our understanding of the critical role of feedback in predator-prey interactions and in feeding relationships that result in community dynamics.

The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.

2. The food supply of the predator population depends entirely on the size of the prey population.

3. The rate of change of population is proportional to its size.

4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.

5. Predators have limitless appetite.

As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey

When multiplied out, the prey equation becomes

dx/dt = αx - βxy

The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt = -

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.

Clone of MAT375 Clone of Bio103 Predator-Prey Model ("Lotka'Volterra")

Jacob Adkins

Building an Ecosystem Model

Axel Caudillo

This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

We incorporate logistic growth into the moose dynamics, and we replace the death flow of the moose with a kill rate modeled from the kill rate data found on the Isle Royale website.

I start with these parameters:
Wolf Death Rate = 0.15
Wolf Birth Rate = 0.0187963
Moose Birth Rate = 0.4
Carrying Capacity = 2000
Initial Moose: 563
Initial Wolves: 20

I used RK-4 with step-size 0.1, from 1959 for 60 years.

The moose birth flow is logistic, MBR*M*(1-M/K)
Moose death flow is Kill Rate (in Moose/Year)
Wolf birth flow is WBR*Kill Rate (in Wolves/Year)
Wolf death flow is WDR*W

Clone of Final Midterm Student version of A More Realistic Model of Isle Royale: Predator Prey Interactions

Jacob Koors

This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

We incorporate logistic growth into the moose dynamics, and we replace the death flow of the moose with a kill rate modeled from the kill rate data found on the Isle Royale website.

I start with these parameters:
Wolf Death Rate = 0.15
Wolf Birth Rate = 0.0187963
Moose Birth Rate = 0.4
Carrying Capacity = 2000
Initial Moose: 563
Initial Wolves: 20

I used RK-4 with step-size 0.1, from 1959 for 60 years.

The moose birth flow is logistic, MBR*M*(1-M/K)
Moose death flow is Kill Rate (in Moose/Year)
Wolf birth flow is WBR*Kill Rate (in Wolves/Year)
Wolf death flow is WDR*W

Clone of Final Midterm Student version of A More Realistic Model of Isle Royale: Predator Prey Interactions

Maria McMahon

Physical meaning of the equations

The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.

2. The food supply of the predator population depends entirely on the size of the prey population.

3. The rate of change of population is proportional to its size.

4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.

5. Predators have limitless appetite.

As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey

When multiplied out, the prey equation becomes

dx/dt = αx - βxy

The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt = -

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.

Clone of Prey&Predator

Eric She

Clone of Bio103 Predator-Prey Model ("Lotka'Volterra")

Tags: Education, Chaos, Ecology, Biology, Population

Thanks to Insight Author: John Petersen

Edits by Andy Long

Everything that follows the dashes was created by John Petersen (or at least came from his Insight model). I just wanted to make a few comments.

We are looking at Hare and Lynx, of course. Clone this insight, and change the names.

Then read the text below, to get acquainted with one of the most important and well-known examples of a simple system of differential equations in all of mathematics.

http://www.nku.edu/~longa/classes/mat375/mathematica/Lotka-Volterra.nb

------------------------------------------------------------

Dynamic simulation modelers are particularly interested in understanding and being able to distinguish between the behavior of stocks and flows that result from internal interactions and those that result from external forces acting on a system.

For some time modelers have been particularly interested in internal interactions that result in stable oscillations in the absence of any external forces acting on a system.

The model in this last scenario was independently developed by Alfred Lotka (1924) and Vito Volterra (1926). Lotka was interested in understanding internal dynamics that might explain oscillations in moth and butterfly populations and the parasitoids that attack them. Volterra was interested in explaining an increase in coastal populations of predatory fish and a decrease in their prey that was observed during World War I when human fishing pressures on the predator species declined.

Both discovered that a relatively simple model is capable of producing the cyclical behaviors they observed.

Since that time, several researchers have been able to reproduce the modeling dynamics in simple experimental systems consisting of only predators and prey. It is now generally recognized that the model world that Lotka and Volterra produced is too simple to explain the complexity of most predator-prey dynamics in nature. And yet, the model significantly advanced our understanding of the critical role of feedback in predator-prey interactions and in feeding relationships that result in community dynamics.

The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.

2. The food supply of the predator population depends entirely on the size of the prey population.

3. The rate of change of population is proportional to its size.

4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.

5. Predators have limitless appetite.

As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey

When multiplied out, the prey equation becomes

dx/dt = αx - βxy

The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt = -

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.

Clone of MAT375 Clone of Bio103 Predator-Prey Model ("Lotka'Volterra")

Jake Moore

STEM-SM combines a simple ecosystem model (modified version of VSEM; Hartig et al. 2019) with a soil moisture model (Guswa et al. (2002) leaky bucket model). Outputs from the soil moisture model influence ecosystem dynamics in three ways.

(1) The ratio of actual transpiration to maximum evapotranspiration (T/ETmax) modifies gross primary productivity (GPP).

(2) Degree of saturation of the soil (Sd) modifies the rate of soil heterotrophic respiration.

(3) Water limitation of GPP (by T/ETmax) and of soil nutrient availability (approximated by Sd) combine with leaf area limitation (approximated by fraction of incident photosynthetically-active radiation that is absorbed) to modify the allocation of net primary productivity to aboveground and belowground parts of the vegetation.

Ecosystem dynamics in turn influence flows of water in to and out of the soil moisture stock. The size of the aboveground biomass stock determines fractional vegetation cover, which modifies interception, soil evaporation and transpiration by plants.

References:

Guswa, A.J., Celia, M.A., Rodriguez-Iturbe, I. (2002) Models of soil moisture dynamics in ecohydrology: a comparative study. Water Resources Research 38, 5-1 - 5-15.

Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools

Clone of Simple Terrestrial Ecosystem Model - Soil Moisture (STEM-SM)

Shudan Xue

last month

Clone of Bio103 Predator-Prey Model ("Lotka'Volterra")

Tags: Education, Chaos, Ecology, Biology, Population

Thanks to Insight Author: John Petersen

Edits by Andy Long

Everything that follows the dashes was created by John Petersen (or at least came from his Insight model). I just wanted to make a few comments.

We are looking at Hare and Lynx, of course. Clone this insight, and change the names.

Then read the text below, to get acquainted with one of the most important and well-known examples of a simple system of differential equations in all of mathematics.

http://www.nku.edu/~longa/classes/mat375/mathematica/Lotka-Volterra.nb

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Dynamic simulation modelers are particularly interested in understanding and being able to distinguish between the behavior of stocks and flows that result from internal interactions and those that result from external forces acting on a system.

For some time modelers have been particularly interested in internal interactions that result in stable oscillations in the absence of any external forces acting on a system.

The model in this last scenario was independently developed by Alfred Lotka (1924) and Vito Volterra (1926). Lotka was interested in understanding internal dynamics that might explain oscillations in moth and butterfly populations and the parasitoids that attack them. Volterra was interested in explaining an increase in coastal populations of predatory fish and a decrease in their prey that was observed during World War I when human fishing pressures on the predator species declined.

Both discovered that a relatively simple model is capable of producing the cyclical behaviors they observed.

Since that time, several researchers have been able to reproduce the modeling dynamics in simple experimental systems consisting of only predators and prey. It is now generally recognized that the model world that Lotka and Volterra produced is too simple to explain the complexity of most predator-prey dynamics in nature. And yet, the model significantly advanced our understanding of the critical role of feedback in predator-prey interactions and in feeding relationships that result in community dynamics.

The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.

2. The food supply of the predator population depends entirely on the size of the prey population.

3. The rate of change of population is proportional to its size.

4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.

5. Predators have limitless appetite.

As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey

When multiplied out, the prey equation becomes

dx/dt = αx - βxy

The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt = -

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.

Clone of MAT375 Clone of Bio103 Predator-Prey Model ("Lotka'Volterra")

Jared Slavey

STEM is a modified implementation of Hartig et al.'s (2019) Very Simple Ecosystem Model (VSEM). The vegetation part of the model has two stocks of biomass carbon (C): aboveground C and belowground C. The soil part of the model has a single stock of soil organic C. Carbon flows into the biomass C stocks via net primary productivity (NPP). Carbon flows out of these stocks and into the soil organic C stock via the loss of aboveground/belowground C through senescence (i.e., abscission of dead leaves and roots). SOC loss is due to heterotrophic respiration of the soil organic matter.

Reference:

Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools

Simple Terrestrial Ecosystem Model (STEM)

Lisa Belyea

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Ecology Models