Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.

This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.

This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.

This model implements the equations proposed by Ketchum in 1954. The rationale behind the concept is that only phytoplankton that grows above a certain rate will not be flushed out of an estuary.

For biological processes:

Pt  =  Po exp(kt)

Where Pt is the phytoplankton biomass at time t, Po is the initial biomass, and k is the growth rate.

For physical processes:

Pm  =  Po (1-r)^m

Where Pm is the phytoplankton biomass after m tidal cycles, and r is the exchange ratio (proportion of estuary water which does not return each tidal cycle).

By substitution, and replacing t by m in the first equation, we get:

Pm = Poexp(km).(1-r)^m

For phytoplankton to exist in an estuary, Pm = Po (at least), i.e. 1 / (1-r)^m = exp(km)
ln(1) - m.ln(1-r) = km
-m.ln(1-r) = km
k = -ln(1-r)

Ketchum (1954) Relation between circulation and planktonic populations in estuaries. Ecology 35: 191-200.

In 2005, Ferreira and co-workers showed that this balance has direct implications on biodiversity of estuarine phytoplankton, and discussed how this could be relevant for water management, in particular for the EU Water Framework Directive 60/2000/EC (Ecological Modelling, 187(4) 513-523).

Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.

Simple model to illustrate a simple simulation of the microalgae biomass production, focusing on the dependent variables such as light, nutrients and other factor that is running for a yearly period.

The biomass model uses an example, Phytoplankton growth based on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Once this is understood, it looks upon the viability of biogas production from the microalgae biomass.

This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.

Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.

Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.

Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.

This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.

Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.

This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.

This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.

This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.

This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.

1D harmful algal bloom model for Alexandrium fundyense based on population dynamics based on models developed for the the Gulf of Maine described in McGillicuddy, Anderson et al. (2005) and model by Stock, McGillicuddy et al. (2005).

This version of these models allows for the Alexandrium cells to control the DIN concentrations and for model limited nutrient scenarios by controlling 'DIN rate in'.  Given that Alexandrium is usually a small component of the phytoplankton biomass in the region, this version of the model is not necessarily appropriate for all events in the Gulf of Maine region and has been developed for regions were Alexandrium populations are the main controlling factor of dissolved nitrogen concentrations.

References:



Stock CA, McGillicuddy DJ, Solow AR, Anderson DA, 2005, Evaluating hypotheses for the initiation and development of Alexandrium  fundyense blooms in the western Gulf of Maine using a coupled physical-biological model. Deep-Sea Research II: Topical Studies in Oceanography, 52(19):2715-2744.

This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.

This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.

Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.

This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.

This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.

This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.