Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator
Insight diagram
This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

Experiment with adjusting the initial number of moose and wolves on the island.
Clone of Isle Royale: Predator Prey Interactions (Español)
Insight diagram
A
Predator-Prey Interactions (Wolf & Moose) QA
Insight diagram
STEM-SM combines a simple ecosystem model (modified version of VSEM; Hartig et al. 2019) with a soil moisture model (Guswa et al. (2002) leaky bucket model). Outputs from the soil moisture model influence ecosystem dynamics in three ways. 
(1) The ratio of actual transpiration to maximum evapotranspiration (T/ETmax) modifies gross primary productivity (GPP).
(2) Degree of saturation of the soil (Sd) modifies the rate of soil heterotrophic respiration.
(3) Water limitation of GPP (by T/ETmax) and of soil nutrient availability (approximated by Sd) combine with leaf area limitation (approximated by fraction of incident photosynthetically-active radiation that is absorbed) to modify the allocation of net primary productivity to aboveground and belowground parts of the vegetation.

Ecosystem dynamics in turn influence flows of water in to and out of the soil moisture stock. The size of the aboveground biomass stock determines fractional vegetation cover, which modifies interception, soil evaporation and transpiration by plants.

References:
Guswa, A.J., Celia, M.A., Rodriguez-Iturbe, I. (2002) Models of soil moisture dynamics in ecohydrology: a comparative study. Water Resources Research 38, 5-1 - 5-15.

Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools

Clone of Simple Terrestrial Ecosystem Model - Soil Moisture (STEM-SM)
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator - 3z MA
Insight diagram
This is an implementation of the 'Very Simple Ecosystem Model' (VSEM) from R package BayesianTools (Hartig et al. 2019). It consists of three stocks: aboveground carbon in plant biomass, belowground carbon in plant biomass and carbon in soil organic matter. 

Reference:
Florian Hartig, Francesco Minunno and Stefan Paul (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools
Clone of Very Simple Ecosystem Model
Insight diagram
STEM-SM combines a simple ecosystem model (modified version of VSEM; Hartig et al. 2019) with a soil moisture model (Guswa et al. (2002) leaky bucket model). Outputs from the soil moisture model influence ecosystem dynamics in three ways. 
(1) The ratio of actual transpiration to maximum evapotranspiration (T/ETmax) modifies gross primary productivity (GPP).
(2) Degree of saturation of the soil (Sd) modifies the rate of soil heterotrophic respiration.
(3) Water limitation of GPP (by T/ETmax) and of soil nutrient availability (approximated by Sd) combine with leaf area limitation (approximated by fraction of incident photosynthetically-active radiation that is absorbed) to modify the allocation of net primary productivity to aboveground and belowground parts of the vegetation.

Ecosystem dynamics in turn influence flows of water in to and out of the soil moisture stock. The size of the aboveground biomass stock determines fractional vegetation cover, which modifies interception, soil evaporation and transpiration by plants.

References:
Guswa, A.J., Celia, M.A., Rodriguez-Iturbe, I. (2002) Models of soil moisture dynamics in ecohydrology: a comparative study. Water Resources Research 38, 5-1 - 5-15.

Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools

Clone of Simple Terrestrial Ecosystem Model - Soil Moisture (STEM-SM)
Insight diagram
This is a model which explains the difference between Mountain bikes riding compared to logging in the Tasmanian forests.
Simulation of Derby Mountain bikes riding versus logging
Insight diagram
This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.


Clone of Key Concepts in Systems Thinking : Predator Prey Interactions
Insight diagram
This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.

Reference:
Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools
Clone of Very Simple Ecosystem Model with Evapotranspiration (VSEM-ET)
Insight diagram
This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale. It was "cloned" from a model that InsightMaker provides to its users, at
https://insightmaker.com/insight/2068/Isle-Royale-Predator-Prey-Interactions
Thanks Scott Fortmann-Roe.

I've created a Mathematica file that replicates the model, at
http://www.nku.edu/~longa/classes/2018spring/mat375/mathematica/Moose-n-Wolf-InsightMaker.nb

It allows one to experiment with adjusting the initial number of moose and wolves on the island.

I used steepest descent in Mathematica to optimize the parameters, with my objective data being the ratio of wolves to moose. You can try my (admittedly) kludgy code, at
http://www.nku.edu/~longa/classes/2018spring/mat375/mathematica/Moose-n-Wolf-InsightMaker-BestFit.nb

{WolfBirthRateFactorStart,
WolfDeathRateStart,
MooseBirthRateStart,
MooseDeathRateFactorStart,
moStart,
woStart} =
{0.000267409,
0.239821,
0.269755,
0.0113679,
591,
23.};

Clone of Isle Royale: Predator/Prey Model for Moose and Wolves
Insight diagram
Start with logistic population dynamics (which can't overshoot) but then add delay in the "feedback signal" (the approach to the carrying capacity). One species, able to exploit one resource, which is available at  a fixed, finite, flow (not a depleting stock). At low populations, growth is exponential. As long as population below carrying capacity, growth continues. Without delay, it will smoothly stabilize at the carrying capacity. But with delay, it will overshoot; but oscillation should dampen, so eventually still stabilizes. Similar dynamics. "from above" (if, e.g., "initial" population somehow above carrying capacity; or, more plausibly, if carrying capacity dynamically falls to some lower level). With more delay, get more extreme overshoot. In "extreme" cases (relatively large delay, large overshoot) we can note asymmetry in "boom" and "bust" - bust is more rapid. This can be interpreted as a very simple version of Bardi's Seneca Cliff.
Population dynamics with overshoot ("Seneca cliff"?)
Insight diagram
This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.

Reference:
Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools
Clone of grassland final final
Insight diagram
This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.

Reference:
Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools
Clone of Very Simple Ecosystem Model with Evapotranspiration (VSEM-ET)
Insight diagram
This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

Experiment with adjusting the initial number of moose and wolves on the island.
Clone of (3) Copy of "Isle Royale: Predator Prey Interactions"
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator
Insight diagram
This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.

Reference:
Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools
Clone of Very Simple Ecosystem Model with Evapotranspiration (VSEM-ET)
Insight diagram

This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

Experiment with adjusting the initial number of moose and wolves on the island.

Clone of Isle Royale: Predator Prey Interactions
Insight diagram
This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

Experiment with adjusting the initial number of moose and wolves on the island.
Clone of Isle Royale: Predator Prey Interactions
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator
Insight diagram
A simple and easy to follow model of how fertility and mortality affect a population, using ferns as an example.
Nick's Fern Population Model
Insight diagram

The purpose of this deer management model is to explore the capacity of wildlife management actions to help us adapt to the effects of climate change.

Clone of Story Telling - Deer Management Under Climate Change