Simple model to illustrate an annual cycle for phytoplankton biomass in temperate waters. Potential primary production uses Steele's equation and a Michaelis-Menten (or Monod) function for nutrient limitation. Respiratory losses are only a function of biomass.
Simple model to illustrate an annual cycle for phytoplankton biomass in temperate waters.
Potential primary production uses Steele's equation and a Michaelis-Menten (or Monod) function for nutrient limitation. Respiratory losses are only a function of biomass.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Simple model to illustrate an annual cycle for phytoplankton biomass in temperate waters. Potential primary production uses Steele's equation and a Michaelis-Menten (or Monod) function for nutrient limitation. Respiratory losses are only a function of biomass.
Simple model to illustrate an annual cycle for phytoplankton biomass in temperate waters.
Potential primary production uses Steele's equation and a Michaelis-Menten (or Monod) function for nutrient limitation. Respiratory losses are only a function of biomass.
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
Simple model to illustrate an annual cycle for phytoplankton biomass in temperate waters. Potential primary production uses Steele's equation and a Michaelis-Menten (or Monod) function for nutrient limitation. Respiratory losses are only a function of biomass.
Simple model to illustrate an annual cycle for phytoplankton biomass in temperate waters.
Potential primary production uses Steele's equation and a Michaelis-Menten (or Monod) function for nutrient limitation. Respiratory losses are only a function of biomass.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Ce modèle est une simulation classique du cycle de productivité dans l'océan, incluant les effets de la thermocline pour désactiver l'advection d'éléments nutritifs dissous et de détritus à la couche superficielle.      Ce modèle illustre un certain nombre de caractéristiques intéressantes notamment
Ce modèle est une simulation classique du cycle de productivité dans l'océan, incluant les effets de la thermocline pour désactiver l'advection d'éléments nutritifs dissous et de détritus à la couche superficielle.  

Ce modèle illustre un certain nombre de caractéristiques intéressantes notamment le lien de trois variables d'état dans un cycle fermé, l'utilisation du temps pour contrôler la durée de l'advection et la fonction modulus pour les données de température qui cyclent annuellement sur plusieurs années. 

Les variables d'état du modèle sont exprimées en unités d'azote (mg N m-3), et l'étalonnage est basé sur:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

Traduction du modèle de Joao G Ferreira (https://insightmaker.com/insight/6838/NPD-model-Nutrients-Phytoplankton-Detritus)
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
 Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I

Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.

Modèle simple pour illustrer la croissance des huîtres sur la base de la production primaire de phytoplancton comme une variable d'état, forcé par la lumière et les éléments nutritifs, en cours d'exécution pour une période annuelle.

La croissance du phytoplancton sur la base de Steele et équations de Michaelis-Menten), où:

Production primaire = (([Pmax] * [I] / [Iopt] * exp (1 - [I] / [Iopt]) * [S]) / ([K] + [S]))

Pmax: production maximale (d-1)
I: L'énergie lumineuse en profondeur de l'intérêt (Ue m-2 s-1)
Iopt: L'énergie lumineuse à laquelle se produit Pmax (Ue m-2 s-1)
S: concentration des éléments nutritifs (N umol L-1)
KS: Demi constants de saturation en nutriments (N umol L-1).

D'autres développements:
- Les éléments nutritifs comme variable d'état dans le cycle de détritus de phytoplancton et d'huîtres de la biomasse.
- Lumière limitée par la concentration de phytoplancton.
- Effet de la température sur le phytoplancton et la croissance des huîtres.