Simple model to illustrate Michaelis-Menten equation for nutrient uptake by phytoplankton.  The equation is:  P = Ppot S / (Ks + S)  Where:  P: Nutrient-limited production (e.g. d-1, or mg C m-2 d-1) Ppot: Potential production (same units as P) S: Nutrient concentation (e.g. umol N L-1) Ks: Half sat
Simple model to illustrate Michaelis-Menten equation for nutrient uptake by phytoplankton.

The equation is:

P = Ppot S / (Ks + S)

Where:

P: Nutrient-limited production (e.g. d-1, or mg C m-2 d-1)
Ppot: Potential production (same units as P)
S: Nutrient concentation (e.g. umol N L-1)
Ks: Half saturation constant for nutrient (same units as S)

The model contains no state variables, just illustrates the rate of production, by making the value of S equal to the timestep (in days). Move the slider to the left for more pronounced hyperbolic response, to the right for linear response.
This model implements the one-dimensional version of the advection-dispersion equation for an estuary. The equation is:  dS/dt = (1/A)d(QS)/dx - (1/A)d(EA)/dx(dS/dx) (Eq. 1)  Where S: salinity (or any other constituent such as chlorophyll or dissolved oxygen), (e.g. kg m-3); t: time (s); A: cross-se
This model implements the one-dimensional version of the advection-dispersion equation for an estuary. The equation is:

dS/dt = (1/A)d(QS)/dx - (1/A)d(EA)/dx(dS/dx) (Eq. 1)

Where S: salinity (or any other constituent such as chlorophyll or dissolved oxygen), (e.g. kg m-3); t: time (s); A: cross-sectional area (m2); Q: river flow (m3 s-1); x: length of box (m); E: dispersion coefficient (m2 s-1).

For a given length delta x, Adx = V, the box volume. For a set value of Q, the equation becomes:

VdS/dt = QdS - (d(EA)/dx) dS (Eq. 2)

EA/x, i.e. (m2 X m2) / (m s) = E(b), the bulk dispersion coefficient, units in m3 s-1, i.e. a flow, equivalent to Q

At steady state, dS/dt = 0, therefore we can rewrite Eq. 2 for one estuarine box as:

Q(Sr-Se)=E(b)r,e(Sr-Se)-E(b)e,s(Se-Ss) (Eq. 3)

Where Sr: river salinity (=0), Se: mean estuary salinity; Ss: mean ocean salinity

E(b)r,e: dispersion coefficient between river and estuary, and E(b)e,s: dispersion coefficient between the estuary and ocean.

By definition the value of E(b)r,e is zero, otherwise we are not at the head (upstream limit of salt intrusion) of the estuary. Likewise Sr is zero, otherwise we're not in the river. Therefore:

QSe=E(b)e,s(Se-Ss) (Eq. 4)

At steady state

E(b)e,s = QSe/(Se-Ss) (Eq 5)

The longitudinal dispersion simulates the turbulent mixiing of water in the estuary during flood and ebb, which supplies salt water to the estuary on the flood tide, and make the sea a little more brackish on the ebb.

You can use the slider to turn off dispersion (set to zero), and see that if the tidal wave did not mix with the estuary water due to turbulence, the estuary would quickly become a freshwater system.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.  We incorporate logistic growth into the moose dynamics, and we replace the death flow of the moose with a kill rate modeled from the kill rate data found on the Isle Royale websi
This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

We incorporate logistic growth into the moose dynamics, and we replace the death flow of the moose with a kill rate modeled from the kill rate data found on the Isle Royale website.

I start with these parameters:
Wolf Death Rate = 0.15
Wolf Birth Rate = 0.0187963
Moose Birth Rate = 0.4
Carrying Capacity = 2000
Initial Moose: 563
Initial Wolves: 20

I used RK-4 with step-size 0.1, from 1959 for 60 years.

The moose birth flow is logistic, MBR*M*(1-M/K)
Moose death flow is Kill Rate (in Moose/Year)
Wolf birth flow is WBR*Kill Rate (in Wolves/Year)
Wolf death flow is WDR*W

 Morecroft, John (2015) Strategic Modelling and Business Dynamics, Ch 1 p13 -14

Morecroft, John (2015) Strategic Modelling and Business Dynamics, Ch 1 p13 -14

Simple model to illustrate Steele's equation for primary production of phytoplankton.  The equation is:  Ppot = Pmax I/Iopt exp(1-I/Iopt)  Where:  Ppot: Potential production (e.g. d-1, or mg C m-2 d-1) Pmax: Maximum production (same units as Ppot) I: Light energy at depth of interest (e.g. uE m-2 s-
Simple model to illustrate Steele's equation for primary production of phytoplankton.

The equation is:

Ppot = Pmax I/Iopt exp(1-I/Iopt)

Where:

Ppot: Potential production (e.g. d-1, or mg C m-2 d-1)
Pmax: Maximum production (same units as Ppot)
I: Light energy at depth of interest (e.g. uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (same units as I)

The model contains no state variables, just illustrates the rate of production, by making the value of I equal to the timestep (in days). Move the slider to the left for more pronounced photoinhibition, to the right for photosaturation.
The Bioresource Model is circular in its nature, and can be divided into two halves, or arcs, within the circle: 1) Primary bioresources = the food and fiber system, and  2) Secondary bioresources = organic waste that is manufactured into secondary bioproducts, e.g. soil amendments (like compost), a
The Bioresource Model is circular in its nature, and can be divided into two halves, or arcs, within the circle:
1) Primary bioresources = the food and fiber system, and
2) Secondary bioresources = organic waste that is manufactured into secondary bioproducts, e.g. soil amendments (like compost), animal feed, materials & chemicals, and energy (fuels, electricity, and CHP)
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.  Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where:   Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]
Simple model to illustrate oyster growth based on primary production of Phytoplankton as a state variable, forced by light and nutrients, running for a yearly period.

Phytoplankton growth based on on Steele's and Michaelis-Menten equations), where: 

Primary Production=(([Pmax]*[I]/[Iopt]*exp(1-[I]/[Iopt])*[S])/([Ks]+[S]))

Pmax: Maximum production (d-1)
I: Light energy at depth of interest (uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (uE m-2 s-1)
S: Nutrient concentration (umol N L-1)
Ks: Half saturation constant for nutrient (umol N L-1).

Further developments:
- Nutrients as state variable in cycle with detritus from phytoplankton and oyster biomass.
- Light limited by the concentration of phytoplankton.
- Temperature effect on phytoplankton and Oyster growth.


Modeling forest succession in a northeast deciduous forest.
Modeling forest succession in a northeast deciduous forest.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.  It illustrates a number of interesting features including the coupling of three state variables in a
This model is a classic simulation of the production cycle in the ocean, including the effects of the thermocline in switching off advection of dissolved nutrients and detritus to the surface layer.

It illustrates a number of interesting features including the coupling of three state variables in a closed cycle, the use of time to control the duration of advection, and the modulus function for cycling annual temperature data over multiple years.

The model state variables are expressed in nitrogen units (mg N m-3), and the calibration is based on:

Baliño, B.M. 1996. Eutrophication of the North Sea, 1980-1990: An evaluation of anthropogenic nutrient inputs using a 2D phytoplankton production model. Dr. scient. thesis, University of Bergen.
 
Fransz, H.G. & Verhagen, J.H.G. 1985. Modelling Research on the Production Cycle of Phytoplankton in the Southern Bight of the Northn Sea in Relation to Riverborne Nutrient Loads. Netherlands Journal of Sea Research 19 (3/4): 241-250.

This model was first implemented in PowerSim some years ago by one of my M.Sc. students, who then went on to become a Buddhist monk. Although this is a very Zen model, as far as I'm aware, the two facts are unrelated.
Simple mass balance model for lakes based on the Vollenweider equation:  dMw/dt = Min - sMw + pMs - Mout  The model was first used in the 1960s to determine the phosphorus concentration in lakes and reservoirs for eutrophication assessment.  This version considers mercury, and adds diagenesis, using
Simple mass balance model for lakes based on the Vollenweider equation:

dMw/dt = Min - sMw + pMs - Mout

The model was first used in the 1960s to determine the phosphorus concentration in lakes and reservoirs for eutrophication assessment.

This version considers mercury, and adds diagenesis, using an extra state variable (mercury in the sediment), and incorporates desorption processes that release mercury trapped in the sediment back to the water column.

The temporal dynamics of the model simulate the typical development of pollution in time.

1. Low loading, low Hg concentration in lake
2. High loading, increasing Hg concentration in lake
3. Desorption rate is low, Hg in sediment increases
4. Measures implemented for source control, loading reduces
5. Hg in lake gradually decreases, but below a certain point, desorption increases, and lake Hg concentration does not improve
6. Recovery only occurs when the secondary load in the sediment is strongly reduced.
Simple model to illustrate Steele's equation for primary production of phytoplankton.  The equation is:  Ppot = Pmax I/Iopt exp(1-I/Iopt)  Where:  Ppot: Potential production (e.g. d-1, or mg C m-2 d-1) Pmax: Maximum production (same units as Ppot) I: Light energy at depth of interest (e.g. uE m-2 s-
Simple model to illustrate Steele's equation for primary production of phytoplankton.

The equation is:

Ppot = Pmax I/Iopt exp(1-I/Iopt)

Where:

Ppot: Potential production (e.g. d-1, or mg C m-2 d-1)
Pmax: Maximum production (same units as Ppot)
I: Light energy at depth of interest (e.g. uE m-2 s-1)
Iopt: Light energy at which Pmax occurs (same units as I)

The model contains no state variables, just illustrates the rate of production, by making the value of I equal to the timestep (in days). Move the slider to the left for more pronounced photoinhibition, to the right for photosaturation.
Dissolved oxygen mass balance in a tide pool, forced by tides and light.
Dissolved oxygen mass balance in a tide pool, forced by tides and light.
Simple model to illustrate an annual cycle for phytoplankton biomass in temperate waters. Potential primary production uses Steele's equation and a Michaelis-Menten (or Monod) function for nutrient limitation. Respiratory losses are only a function of biomass.
Simple model to illustrate an annual cycle for phytoplankton biomass in temperate waters.
Potential primary production uses Steele's equation and a Michaelis-Menten (or Monod) function for nutrient limitation. Respiratory losses are only a function of biomass.